Peristome development pattern in Encalypta poses a problem: what is the primary peristomial layer in mosses?
Michael S. Ignatov, Ulyana N. Spirina, Maria A. Kolesnikova, Darya A. Ashikhmina, Elena A. Ignatova, Svetlana V. Polevova

Literature Cited
Blomquist,H.L. & L.L. Robertson. . 1941 . The development of the peristome in Aulacomnium heterostichum. Bulletin of the Torrey Botanical Club 65: 569–584.Google Scholar, Crossref
Brotherus,V.F. . 1924 . Musci. In: Engler, A. & K. Prantl (eds.) Die Natürlichen Pflanzenfamilien. 2, 10. Berlin: Duncker and Humblot: 143–478.Google Scholar
Budke,J.M., C.S. Jones & B. Goffinet. 2007. Development of the enigmatic peristome of Timmia megapolitana (Timmiaceae; Bryophyta). American Journal of Botany 94(3): 460–467.Google Scholar, Crossref
Cox,C.J., B. Goffinet, N.J. Wickett, S.B. Boles & A.J. Shaw. 2010. Moss diversity: a molecular phylogenetic analysis of genera. Phytotaxa 9: 175–195.Google Scholar, Crossref
Edwards,S.R. 1979. Taxonomic implications of cell patterns in haplolepideous moss peristomes. In: Clarke, G.C.S. & J.G. Duckett (eds.). Bryophyte systematics. Academic Press, London, UK.: 317–346.Google Scholar
Edwards,S.R. 1984. Homologies and inter-relations of moss peristomes. In: Schuster, R.M. (ed.) New manual of bryology, vol. 2. Hattori Botanical Laboratory, Nichinan, Japan: 658–695.Google Scholar
Evans,A.W. & H.D. Hooker, jr. 1913. Development of the peristome in Ceratodon purpureus. Bulletin of the Torrey Botanical Club 40: 97–109.Google Scholar, Crossref
FEDOSOV,V.E., A.V. FEDOROVA, A.E. FEDOSOV & M.S. IGNATOV. 2016. Phylogenetic inference and peristome evolution in haplolepideous mosses, focusing on Pseudoditrichaceae and Ditrichaceae s.l. Botanical Journal of the Linnean Society 181: 139–155.Google Scholar, Crossref
Frey,W. & M. Stech. 2009. Bryophyta (Musci, mosses). In: Frey, W. (ed.). Syllabus of plant families A. Engler’s Syllabus der Pflanzenfamilien. Part 3. Bryophytes and seedless vascular plants. 13th ed. Gebr. Borntraeger Verlagsbuchhandlung, Stuttgart, Germany: 116–257.Google Scholar
Goffinet,B., W.R. Buck & A.J. Shaw. 2009. Morphology, anatomy, and classification of the Bryophyta. In: Goffinet, B. & A.J. Shaw (eds.). Bryophyte biology, 2nd edn. Cambridge: Cambridge University Press: 55–138.Google Scholar
Goffinet,B., J. Shaw, L.E. Anderson & B.D. Mishler. 1999. Peristome development in mosses in relation to systematics and evolution. V. Diplolepideae: Orthotrichaceae. Bryologist 102: 581–594.Google Scholar, Crossref
Haeckel,E. 1866. Generelle Morphologie der Organismen : allgemeine Grundzüge der organischen Formen-Wissenschaft, mechanisch begründet durch die von Charles Darwin reformirete Descendenz-Theorie. Berlin.Google Scholar
HORTON,D.G. 1982. A revision of the Encalyptaceae (Musci) with particular reference to the north American taxa Part I. Journal of the Hattori Botanical Laboratory 53: 365–418.Google Scholar
Horton,D.G. 1983. A revision of the Encalyptaceae (Musci) with particular reference to the north American taxa Part II. Journal of the Hattori Botanical Laboratory 54: 353–532.Google Scholar
Ignatov,M.S., U.N. Spirina, E.A. Ignatova, M. Krug & D. Quandt. 2015. On the systematic position of the moss genus Catoscopium, with a new approach to the peristome reduction study. Arctoa 24(2): 389-415.Google Scholar, Crossref
Ignatov,M.S., V.E. Fedosov, A.V. Fedorova & E.A. Ignatova. 2016. On the systematic position of Discelium (Bryophyta). Arctoa 25(2): 278–284.Google Scholar, Crossref
Kienitz-Gerloff,F. 1878. Untersuchungen uber die Entwick-lungsgeschichte der Laubmooskapsel und die Embryoen- twicklung einiger Polypodiaceen. Botanische Zeitung 36: 33–64.Google Scholar
Kreulen,S.J.W. 1972. Features of single- and double-peristome capsules. Homology of layers and ontogeny of outer spore sac. Lindbergia 1: 153–160.Google Scholar
Liu,Y., J.M. Budke & B. Goffinet. 2012. Phylogenetic inference rejects sporophyte based classification of the Funariaceae (Bryophyta): rapid radiation suggests rampant homoplasy in sporophyte evolution. Molecular Phylogenetics and Evolution 62: 130–145. doi:10.1016/j.ympev.2011.09.010.Google Scholar, Crossref
Loeske,L. 1929. Funariaceae. Die Laubmoose Europas (Loeske), Bd. 2. Max Lande, Berlin, 120 pp.Google Scholar
Medina,R., M. Johnson, Y. Liu, N. Wilding, T. A. J. Hedderson, N. Wickett & B. Goffinet. 2018. Evolutionary dynamism in bryophytes: Phylogenomic inferences confirm rapid radiation in the moss family Funariaceae. Molecular Phylogenetics and Evolution 120: 240–247.Google Scholar, Crossref
MUELLER,D. M. J. 1973. The peristome of Fissidens limbatus Sullivant. University of California Publications in Botany 63: 1–34.Google Scholar
Neesvon Esenbeck, C. G. D., C. F. Hornschuch & J. J. W. Sturm. 1827. Bryologia Germanica. Bd. 2(1). Friedrich Fleischer, Leipzig, 182 pp.Google Scholar
Newton,A.E., C.J. Cox, J.G. Duckett, J.A. Wheeler, B. Goffinet, T.A.J. Hedderson & B.D. Mishler. 2000. Evolution of the major moss lineages: phylogenetic analyses based on multiple gene sequences and morphology. Bryologist 103: 187–211.Google Scholar, Crossref
Nyholm,E. 1998. Illustrated flora of Nordic Mosses. Fasc. 4. Aulacomniaceae Meesiaceae – Catoscopiaceae – Bartramiaceae – Timmiaceae – Encalyptaceae – Grimmiaceae – Ptychomitriaceae – Hedwigiaceae – Orthotrichaceae. – Nordic Bryological Society, Copenhagen & Lund, pp. 249–405.Google Scholar
Philibert,H. 1889. Etude sur la péristome. Huitiиme article. Difference entre les Nématodontées et les Arthrodontéees; transition entre ces deux groupes (Suite). Revue Bryologique 16: 1–9, 39–44, 67–77.Google Scholar
Saito,S. 1956. Studies on the development of the peristome in Musci II. On the peristome in Dicranum japonicum Mitt. Botanical Magazine, Tokyo 69(812): 53–59.Google Scholar
Saito,S. & S. Shimoze. 1955. Studies on the development of the peristome in Musci I. On the peristome in Bartramia crispula Schimp. Botanical Magazine, Tokyo 69(812): 53–59.Google Scholar
Schwartz,O.M. . 1994 . The development of the peristome-forming layers in the Funariaceae. International Journal of Plant Science 155: 640–657.Google Scholar, Crossref
Schimper,W.P. 1876. Synopsis Muscorum Europeorum. 2d ed. E. Schweizerbart, Stuttgart, 866 pp.Google Scholar
Shaw,A.J., L.E. Anderson & B.D. Mishler. . 1987 . Peristome development in mosses in relation to systematics and evolution 1. Diphyscium foliosum (Diphysciaceae). Memoirs of the New York Botanical Garden 45: 55–70.Google Scholar
Shaw,A.J., L.E. Anderson & B.D. Mishler. . 1989. a. Peristome development in mosses in relation to systematics and evolution III. Funaria hygrometrica, Bryum pseudocapillare, and B. bicolor. Systematic Botany 14: 24–36.Google Scholar
Shaw,A.J., L.E. Anderson & B.D. Mishler. . 1989. b. Peristome development in mosses in relation to systematics and evolution IV. Haplolepideae: Ditrichaceae and Dicranaceae. Bryologist 92: 314–323.Google Scholar, Crossref
Shaw,J. & K. Renzaglia. 2004. Phylogeny and diversification of Bryophytes. American Journal of Botany 91(10): 1557–1581.Google Scholar, Crossref
Shaw,A.J. & H. Robinson. . 1984 . On the development, evolution, and function of peristome in mosses. Journal of the Hattori Botanical Laboratory 57: 319–335.Google Scholar
Shaw,A.J., P. Szövényi & B. Shaw. 2011. Bryophyte diversity and evolution: windows into the early evolution of land plants. American Journal of Botany 98(3): 352–369.Google Scholar, Crossref
TAYLOR,E. C. 1962. The Philibert peristome articles. An abridged translation. Bryologist 65: 175–212.Google Scholar, Crossref
Tsubota,H., E. De Luna, D. González, M.S. Ignatov & H. Deguchi. 2004. Molecular phylogenetic and ordinal relationships based on analyses of a lage-scale datd set of 600 rbcL sequences of mosses. Hikobia 14: 149–170.Google Scholar
YOUNG,R.L. G.S. BEVER, Z. WANG & G.P. WAGNER. 2011. Identity of the avian wing digits: Problems resolved and unsolved. Developmental Dynamics 240: 1042–1053.Google Scholar, Crossref
ZHU,J. & S. MACKEM. 2013. a. Analysis of mutants with altered shh activity and posterior digit loss supports a biphasic model for shh function as a morphogen and mitogen. Developmental Dynamics 240(5): 1303-1310. DOI: https://doi.org/10.1002/dvdy.22637Google Scholar
ZHU,J. & S. MACKEM. 2013. b. Tracing the Evolution of Avian Wing Digits. Current Biology 23(12): R538-R544.Google Scholar, Crossref