Saaristo M.I. 2007. A new subfamily of linyphiid spiders based on a new genus created for the keyserlingi-group of the genus Lepthyphantes (Aranei: Linyphiidae) // Arthropoda Selecta. Vol.16. No.1: 33–42 [in English].
Zoological Museum, Centre for Biodiversity, University of Turku, FI-20014 Turku Finland (deceased).
Key words: Micronetinae, Linyphiinae, taxonomy, new combination, new synonym, male palp, epigyne.
Abstract. A new linyphiid subfamily, Ipainae subfam.n., is established to include seven genera and 26 species, of which one genus is new to science: (1) The type genus Ipa gen.n., containing I. keyserlingi (Ausserer, 1867), comb.n. (the type species), I. terrenus (L. Koch, 1879), comb.n. = I. quadrimaculatus (Kulczy?ski, 1898), syn.n., comb.n., I. spasskyi (Tanasevitch, 1986), comb.n. and I. pepticus (Tanasevitch, 1988), comb.n., all ex Lepthypahntes; (2) Epibellowia Tanasevitch, 1996, containing E. septentrionalis (Oi, 1960) (the type species), E. pacificus (Eskov et Marusik, 1992) and E. enormitus (Tanasevitch, 1988); (3) Metalepthyphantes Locket, 1968 ad partem, containing M. bifoliatus Locket, 1968, M. carinatus Locket, 1968, M. clavator Locket, 1968, M. dentiferens Bosmans, 1979, M. machadoi Locket, 1968 (the type species), M. ovatus Scharff, 1990, M. perexiguus (Simon et Fage, 1922), M. praecipuus Locket, 1968 and M. vicinus Locket, 1968; (4) Solenysa Simon, 1894, containing S. circularis Gao, Zhu et Sha, 1993, S. geumoensis Seo, 1996, S. longqiensis Li et Song, 1992, S. melloteei Simon, 1894 (the type species), S. protrudens Gao, Zhu et Sha, 1993 and S. wulingensis Li et Song, 1992; (5) Uralophantes Esynin, 1993, containing U. troitskensis Esynin, 1993 (the type species); (6) Wubanoides Eskov, 1986, containing W. uralensis (Pakhorukov, 1981) = W. longicornis Eskov, 1986 (the type species) and W. fissus (Kulczy?ski, 1926); and (7) Epigytholus Tanasevitch, 1996, containing E. tuvensis Tanasevitch, 1996 (the type species). Because both Micronetinae and some representatives of the new subfamily Ipainae have the so-called Fickert’s gland inside their radix, they are hypothesized to be sister groups despite otherwise greatly deviating secondary genital organs.