Shcherbakov D.E. 2021. New Curvicubitidae and Paraknightiidae (Homoptera: Cicadomorpha) from the Triassic of Central Asia // Russian Entomol. J. Vol.30. No.2: 129–134 [in English].

Borissiak Paleontological Institute, Russian Academy of Sciences, Profsoyuznaya St. 123, Moscow 117647, Russia. E-mail:

doi: 10.15298/rusentj.30.2.02

ABSTRACT. Three new species and one new genus of Cicadomorpha are described from the Middle or Upper Triassic of Madygen, Kyrgyzstan. The family Curvicubitidae includes one genus, Beaconiella Evans, 1963 (= Curvicubitus Hong, 1984, syn.n.; Beaconiella triassica (Hong, 1984), comb.n.). The wing venation of Beaconiella pulchra sp.n. and closely related B. cincta sp.n. confirms that this family belongs to Pereborioidea, despite some similarities with Palaeontinoidea. Triknightia mira gen. et sp.n. (Scytinopteroidea: Paraknightiidae) combines the heteropterous forewing structure and folding with the homopterous head structure. This is the first Mesozoic find of Paraknightiidae, representing an early side branch near the root of true Heteroptera. The head structure is first described for paraknightiids, and the clavus and hindwing for curvicubitids — their hindwing bears a row of hooks instead of coupling lobe. Curvicubitidae and Triknightia gen.n. are the last relics of the Permian groups Pereborioidea and Paraknightiidae in the Triassic fauna, and both taxa are highly advanced compared to their Permian relatives.

KEY WORDS: Hemiptera, Auchenorrhyncha, Pereborioidea, Palaeontinoidea, Scytinopteroidea, Heteroptera, true bugs, forewing, hemelytron, venation, nodal line, phylogeny, fossil, relict, Lagerstätte, Madygen.

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