Scientific peer-reviewed journal INVERTEBRATE ZOOLOGY publishes original papers, reviews and brief communications on morphology, anatomy, embryology, taxonomy, phylogeny, and ecology of any group of invertebrates from protistans to lower chordates. Invertebrate Zoo­logy accepts manuscripts in both English and Russian and publishes them in two identical versions, printed and electronic. Papers dealing with taxonomic revisions are available online upon printing out to avoid any conflicts about priority issues.

In addition to the rules given herein, please examine Template.


  • English language editing

Authors are responsible for submitting papers written in grammatically correct English language of sufficient quality. Academic style should be used for all types of manuscripts. Authors are advised to use text editing services provided by either professional organisations or their native speaking colleagues. Please make sure you acknowledge their help in a particular section of your manuscript. Translation of the Title, Abstract and Figure captions into Russian is provided by the editors.


  • Structure of the manuscript

Title page

The title of the paper should be concise and informative.


Please include the following author information:

The names, affiliations and ORCID numbers of all the authors, including institution, city, country; e-mail address of the corresponding author


  • (up to 500 words) should not contain any undefined abbreviations or unspecified references


Provide 4–7 keywords. Please avoid abbreviations except widely accepted in the field.


Main text body should contain


  • . Please provide clear objectives of the work and avoid summary of the results.


Material and methods.

Provide sufficient detail to allow the work to be reproduced. References to already published methods should be provided with the descriptions of modifications.


  • should be clear and concise


A combined Results and Discussion section is sometimes appropriate. Avoid extensive citations and discussion of published literature.


  • The main conclusions of the study may be presented in a short Conclusions section, which alternatively may be included into Discussion or Results and Discussion section.


Acknowledgements of people, grants, funds, etc. should be provided as a separate section. The names of funding organizations should be written in full.


Reference list

The list of references should only include works that are cited in the text. Reference list entries should be alphabetized by the last names of the first author of each work. Please use alphabetic order according to the following rules: 1) For the references with one author, by the name of the author, then chronologically; 2) For the references with two authors, by the name of the first author, then the name of the second author, then chronologically; 3) For the references with more than two authors, by the name of the first author, then chronologically.

Titles of Russian papers (or other papers using non-Latin alphabets) should be translated while the titles of the journals or books should be transliterated (e.g. Trudy Instituta Okeanologii) in the list of references. Original language should be indicated in square brackets at the end of the reference, e.g. [in Russian].

References should be cited in the text by name and year, separated by comma in round brackets, e.g. (Huys, Boxshall, 1991), (Huys, Boxshall, 1991; Tunnicliffe et al., 1998). The references to the papers published in Russian should be transliterated into Latin letters. A paper with more than two authors should be cited as (Makarov et al., 1982)

If available, please always include DOIs as full DOI links in your reference list (e.g. “”)


Reference to a journal article:

Tunnicliffe V., McArthur A.G., McHugh D. 1998. A biogeographical perspective of the deep-sea hydrothermal vent fauna // Adv. Mar. Biol. Vol.34. P.355–442.

Marusik Yu.M. 1987. [Three new species of the family Nesticidae (Aranei) from the fauna of the USSR] // Zool. Zhurn. Vol.66. No.3. P.461–463 [in Russian, with English summary].

Reference to a book:

Hoffman R.L. 1980 (for 1979). Classification of the Diplopoda. Mus. hist. nat. Geneve. 237 p.

Reference to a chapter in an edited book:

Mironov A.N., Gebruk A.V., Moskalev L.I. 2002. Biogeography of hydrothermal vent communities and obligate hydrothermal taxa // A. Gebruk (ed.). Biology of hydrothermal systems. Moscow: KMK Sci. Press Ltd. P.410–455.

Reference to a dissertation:

Kovblyuk M.M. 2006. [Gnaphosid spiders (Arachnida: Aranei) in Crimean fauna]. [Thesis of Candidate (Ph.D.) of Biological Sci. Degree]. Kyiv: Institute of Zoology Ukrainian Acad. of Sci. 200 p. [In Ukrainian, with Russian and English summary]

Figure captions should be included in the text file of the manuscript, not in the figure file. Figure captions should begin with the term Fig. followed by the figure number. All elements found in the figure plate should be identified in the figure caption; previously published material must be identified by providing the reference citation at the end of the figure caption. Abbreviations must be provided in alphabetical order. 

Diagrams and graphs should be scanned in black-and-white (Bitmap) mode.

The abbreviations for figures must be performed by usual (NOT italics, NOT bold, NOT underlined) small LATIN letters, in Times New Roman, size 14.

The figure captions should be submitted on the separate sheet and must contain:

Figure number. General title of the figure applying to ALL its parts. Part number — description of the photograph (picture); The next part number — description of the photograph (picture).

Abbreviations (in alphabetical order): ab — …….; bs — ……. Scale bar …….. mm (or Scale bars: А — …; B — …

The general title of figure should be separated from the number of its first part by a point. The number of a part (Capital Latin letter) should be separated from the describing by a long dash («Insert – Symbol – choose the symbol – Paste»). The same long dash is used in abbreviations and scales.

The descriptions of the figures parts should be separated by semicolons. And the same is used for the list of abbreviations and scales.

Please follow the formatting of the figure captions as in the example below:

Fig. 1. Cells of exocyst of Aploparaksis bulbocirrus at the stage of late sclexogenesis. A–B — sagittal sections; C — transversal section.

A — muscle cells and tegument’s cyton of internal side of exocyst; B — cells in caudal part of exocyst; C — cell with signs of undifferentiated and “dark” cells in caudal part of exocyst.

Abbreviations: dac — “dark cell”; dc — distal part of tegument; ger — granular endoplasmic reticulum; m — mitochondrion; mc — muscle cells; mi — microvillus; n — nucleus; nu — nucleolus. Scale bar: A — 2 µm; B, C — 2.5 µm.


  • Text formatting

Manuscripts should be submitted in Word. Save your file in doc format.

Normal, plain font (Times New Roman, font size 12, interline spacing 1.5) should be used for the text without any specific formatting.

Generic and species names must be italicized.

Please use automatic page numbering function to number the pages.

  • Illustrations

All illustrations should be provided in TIFF format as separate files.

Figure files should be named as «AuthorSurname_Figure_Number». For example «Temereva_Figure_01»

The maximal size of figure table is 13 cm width and 17 (or less) cm height. The size of single column in the table is 6.5 cm width and 17 (or less) cm height.

Halftones an color figures should have a minimum resolution of 300 dpi; 600 dpi – for diagrams and graphs. Color illustrations should be submitted as RGB (8 bits per channel).

The parts of figure table (separate pictures) must be numbered by capital latin letters. Parts of figures and abbreviations should be done as Arial, font size 14 or 12.


  • Supplementary materials

Supplementary electronic materials can be submitted to be published on the website after acceptance of the manuscript without any conversion, editing, or reformatting.

All supplementary materials, must have a citation in the manuscript, similar to that of figures and tables. (Supplementary file 1)

A concise caption describing the content of the file should be provided for each supplementary material.


  • How to submit

To submit your manuscript, please email us ( separately attached text and illustrations files.

Authors are encouraged to provide names and contact details of three potential referees.


  • Cost of color figures

Online publication of color illustrations is free of charge.

For color illustration in the printed version, authors will have to pay 100 EUR per article, irrespective of the number of figures.